STATE COLLEGE — ESPN analyst Kirk Herbstreit was just one of many.

Doubters predicted that Penn State, led by former walk-on Matt McGloin, would have trouble moving the ball against Michigan.

Herbstreit emphasized that point Saturday morning on ESPN’s College Gameday, which must have irritated the redshirt sophomore.

His response? McGloin completed 17 of 28 pass attempts for 250 yards and a touchdown. He ran for a touchdown, too.

On the field immediately following the game, McGloin called out Herbstreit, saying he was “gunning for him,” trying to prove the analyst wrong.

“I was a little upset this morning (when) I watched it,” McGloin said after the game. “He says we’re not going to be able to move the ball today? I think we moved it pretty good.”

Pretty good may be an understatement.

The Nittany Lions posted 435 yards of total offense, making it the fifth time this season the Wolverine defense has given up more than 400 total yards.

“As a quarterback, you have to take what the defense gives you, and I think I did that well tonight,” McGloin said.

McGloin said Michigan did a few things differently from what he had seen on film last week, but he felt prepared for the Wolverine defense overall.

The quarterback started Saturday’s game because the Nittany Lions’ regular starter, Rob Bolden, was knocked out of last weekend’s game with a concussion. McGloin is Penn State’s third-string quarterback, but he performed above expectations on Saturday.

“When you’re playing someone like Michigan, who has someone like Denard Robinson, the object of the game is to keep him off the field,” McGloin said.

Royster’s record day: On the second play of Penn State’s first possession Saturday night, Nittany Lion senior running back Evan Royster dashed for 20 yards.

On that play, Royster became Penn State’s all-time leading rusher. He ended the night with 3,518 yards, breaking Curt Warner’s mark of 3,398 yards, which had stood since 1982.

Royster’s 150 yards and two touchdowns on the ground complemented McGloin’s passing attack well, making it easy for the Nittany Lions to build up an early lead against the Wolverines.

“Evan had a great night, congratulations to him,” McGloin said. “He really had a breakout game. That really set up the passes downfield. We went play action a lot tonight. Because of Evan’s running, we were able to do that.”

Even longtime Penn State coach Joe Paterno, who has seen plenty of running backs in blue and white over the years, had praise for Royster.

“That’s the best he’s played,” Paterno said. “He’s a good back, a good solid back. But he’s like all of them — he needs some running room. I think he got some today, and he took advantage of it.”

Hagerup kicking? Redshirt freshman kicker Seth Broekhuizen kicked off to Penn State after Michigan’s first touchdown of the game, but the rest of the game, true freshman punter Will Hagerup assumed all Wolverine kickoff duties.

“Seth is struggling with that a little bit, and we practiced (Hagerup) quite a bit this week and he was doing pretty good,” Michigan coach Rich Rodriguez said after the game. “He kicked pretty well today.”

Hagerup said he was told prior to Michigan’s second kickoff that the responsibility would be his. He said though he practiced during the week, he had “no idea” whether he’d actually handle a kickoff.

“I think I did alright,” Hagerup said. “It’s like anything. I’ve been practicing a lot this year. You’re going to be rusty, but I can definitely get better at it.”

Hagerup said he expects to compete for the job this week in practice.

Injury notes: Nose tackle Mike Martin injured his right ankle two weeks ago against Iowa, and in the first half Saturday, he hurt his left one.

“That’s a blow because Mike makes a difference when he’s in there,” Rodriguez said.

Martin told the Daily he would "be okay."

Cornerback J.T. Floyd, whom Rodriguez said suffered a stinger, left and re-entered the game.

Rodriguez also said Robinson’s shoulder, which has been banged up in recent weeks, is still not 100 percent healthy. When asked about it, Robinson said he felt good.